(Bovidae)**
* * The family of bovids, or bulls, is the largest and most diverse group of artiodactyls, including 45-50 modern genera and about 130 species.
* * * Due to seasonal uneven growth, “annual rings” are visible on the horns of some artiodactyls, allowing one to determine the age of the animal.
* Unlike deer, bovids never have upper canines; the molars have a higher crown and a more complex chewing surface. The number of fingers is sometimes reduced to two.
* * Being numerous and diverse in Africa and Eurasia, bovids in limited numbers penetrated into North America only in the Pleistocene, crossing the Beringian land. Now only 5-6 species from 4 genera (subfamilies Caprinae and Bovinae) are found here. Bovids never reached South America and Australia, as well as many islands and archipelagos. In Russia 12 wild species bovids from 8 genera.
* The range of the bison in the Middle Bek covered Central and Eastern Europe from Germany and Hungary to the Don basin and the Caucasus. Wild bison survived for the longest time in the Caucasus and in the area of Belovezhskaya Pushcha. By the 20s of the 20th century it disappeared from nature. 45 animals of the nominate subspecies are preserved in zoos, mainly in Poland. As a result of restoration work in captivity (including absorptive crossing with bison), the bison was saved as a species and returned to some of its former habitats in nature reserves in Poland and former USSR. In the Moscow region there is a nursery at the Prioksko-Terrasny Nature Reserve; groups of bison have been reacclimatized in the Caucasus, in the Carpathians, and brought to the Tien Shan. Now there are about 1.5 thousand animals in the world.
* The height of bulls at the withers is up to 2 m, weight is up to 850 kg. Cows can be half the weight.
* * The bison differs from the bison in its larger size. but at the same time with a lighter physique. He doesn't have that much big head, set much higher, longer and thinner horns, arched back profile, more developed rear part of the body. The legs are noticeably higher, the tail is longer. The coat is more uniform in length and a uniform brown color. In the diet higher value branches and leaves play (in total, bison consume more than 200 plants).
* The last bison of a special Caucasian subspecies (B. b. caucasicus), distinguished by very dark curly hair and some other features, were exterminated by poachers in 1925 - 1927 in the Teberda region. Now in Teberdi and other parks and reserves of the Caucasus live herds of bison, Belovezhsky in origin, as well as bison. Sedentary on the plains, in the mountains bison make vertical migrations, rising in the summer to 2000 m above sea level.
* Bison can jump up to 3 m in length and up to 2 m in height.
* During the rut, the bull has a “harem” of 2-6 females with him.
* * The weight of a newborn is about 22 kg, lactation lasts 5-6 months (sometimes up to a year), but the calf begins to eat grass from 2-3 weeks. Sometimes the calf remains with its mother for up to 2 years, despite the fact that, under favorable conditions, the female brings new offspring next spring. Puberty occurs at 1.5-2 years, but animals reach their final size by 5-8 years. Maximum life expectancy is about 40 years.
* Bison entered America from Eurasia in glacial period. Their different forms will replace each other in the tundra-steppes, forests and prairies; some species were much larger than the modern steppe bison (Bison bison); the span of their long horns reached 2 meters or more. Despite the common American name (Buffalo), bison, like bison, are more closely related to bulls than to buffalos.
The number of surviving bison reached, according to the exact information of William Gornedey, on January 1, 1889, up to 835 animals, including those 200 bulls that live under government protection in Yellowstone Park. This extermination of bison began in the seventies, when the railroads were built
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Akimushkin Igor Ivanovich (1929-1993)
Born in Moscow in the family of an engineer. Graduated from the Faculty of Biology and Soil Sciences of Moscow State University (1952). Published since 1956.
His first books for children appeared in 1961: “Traces of Unseen Beasts” and “The Path of Legends: Tales of Unicorns and Basilisks.”
Igor Ivanovich wrote a number of books for children, using techniques that are typical for fairy tales and travel. These are: “Once upon a time there was a squirrel”, “Once upon a time there was a beaver”, “Once upon a time there was a hedgehog”, “Building animals”, “Who flies without wings?”, “Different animals”, “Why is a rabbit not like a hare” and etc.
For teenagers, Akimushkin wrote books of a more complex genre - encyclopedic ones: “River and Sea Animals”, “Entertaining Biology”, “The Vanished World”, “The Tragedy of Wild Animals”, etc.
Akimushkin’s focus is on current issues of development, conservation and study of the animal world, research on the behavior and psyche of animals. He wrote not only books for children and youth; but also scripts for popular science films. A number of Akimushkin’s works have been translated into foreign languages. His most famous work is the book “Animal World”.
“The World of Animals” is the most famous work of Igor Ivanovich Akimushkin, which has gone through several reprints. They summarize a huge amount of scientific material, use a more modern classification scheme for the animal world, many different facts from the life of animals, birds, fish, insects and reptiles, beautiful illustrations, photographs, funny stories and legends, incidents from life and notes from an observer-naturalist. Six volumes of “The World of Animals” by Igor Ivanovich Akimushkin were published one after another over the course of a decade - from 1971 to 1981. They were published by the publishing house “Young Guard” in the popular “Eureka” series. In ten years, readers have managed to grow up and fall in love with these books for the rest of their lives. The first and second talked about mammals, the third - about birds, the fourth - about fish, amphibians and reptiles, the fifth - about insects, the sixth - about domestic animals.
The first book, “The World of Animals,” tells about seven orders of mammals: cloacals, marsupials, insectivores, woolly wings, carnivores, equids and artiodactyls.
Why was Australia inhabited only by marsupials and egg-laying animals before the arrival of humans? Who is stronger: a lion, a tiger or a bear? Secrets behind the needles - about the incomprehensible habits of hedgehogs. Igor Akimushkin invites readers to take him on a fascinating journey into the animal kingdom. In this book, the author talks about the world of mammals. The theme of human responsibility for the fate of animals on our planet runs through the entire book.
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In bovids, both males and females (with rare exceptions) wear a pair, or even two pairs of horns. The fact that their horns are hollow, that is, empty inside, seems to be beyond doubt, and, however, this is not entirely true: the horns seem to be “attached” to rods protruding from the frontal bone.
Shape and size? Here, as writers of old used to say, “the pen falls out of your hands.” Lumpy, folded, faceted, smooth, twisted, twisted, just straight - in general, all sorts. The length and width are also different: from miniature stilettos to huge rapiers. The girth of the argali horns at the base, for example, is about 50 centimeters.
The horns of bovids grow throughout their lives, but never branch. They consist of a substance of epidermal origin, an excellent material for making glue (the Chinese, as usual, also make medicines from them). Strongly civilized hunters (for example, those who have impoverished the fauna of Africa) use hollow horns for... Well, E. Hemingway answered this question to one African: “Tell him that, according to the customs of our tribe, we give horns to our richest friends. Also say that this is a very exciting event and sometimes some of our fellow tribesmen are chased by people with unloaded pistols.”
Some zoologists call bovid animals “horned.” Everyone has horns. All sorts of horns: straight and sharp meter bayonets; curved like sabers, twisted into a corkscrew; twisted into a “ram’s horn”; small, like stilettos, - great variety. Horns are found in females and males, less often only in males. Some will be born with the beginnings of horns, many are polled at birth.
Why are horns needed? It would seem an idle question: for defense and attack. We always thought so. But in Lately doubts arose.
If for defense, then why do females, who in this case need horns most of all, often have none at all or are small? Previously, it was self-evident that females with cubs were protected by strong and horned males. But the males of many bovids do not even think about protecting their females and children. If the predator is strong and there is no point in fighting, they usually run away first. But even if the predator is small and the horns could be useful to drive it away, even such strange things have been noticed at first glance: the male rushes not to help the female, but at her! When, for example, a female Thomson's gazelle happens to wound and drive a jackal away from her cub and she rushes in pursuit of the predator, the male immediately rushes after her and forces her to turn back. For what? Yes, because he is afraid that she will run away from his harem. This possessive – or rather, sexual – instinct suppresses the male’s instinct to care for the offspring.
Not everyone does this, but many do. True, among musk oxen and American snow goats, when threatened by a wolf attack, the males always join forces to repel predators. Large bulls, buffaloes for example, do not give in to lions. It's right. But here’s what’s interesting: buffaloes, musk oxen, and snow goats, that is, those who use their horns most actively, do not have the best structure at all. Or small, like snow goat, or too curved. And here we would need straight ones, sharp as swords.
But maybe horns are necessary to fight with relatives for females and territory? Indeed, male gazelles, for example, and many other bovids butt heads with each other ten times a day. But the horns are used with great care, not for mutilation, but for ritual confrontation. Of course, it happens, and often, when mortal wounds are inflicted by a blow to the side, in the most unprotected place. But this is rather an exception. Usually, before fighting, males, according to the rules that evolution has laid down in their instincts, stand in a certain position: head to head. Here the blows are delivered flat with the horns. Such fencing, no better word needed, is customary among antelopes. At the same time, some even kneel (roan antelopes and nilgai) and, straining their strength, try to push away or knock down the enemy. Roan antelopes rest in this power struggle with the middle of their back-curved horns, and nilgai with their foreheads. Nilgai, intertwining their necks, try to knock down their opponent. And all this while on my knees!
By the way, neck wrestling is one of the original ritual forms. Just like bites. Over the course of evolution, in many species it was replaced by fencing and fighting with locked horns. It is interesting that in females and cubs that do not have horns or they are small, more ancient ritual fighting tactics have been preserved as a kind of atavism: biting, kicking, clasping the neck, hitting the side with the forehead.
It is the hornless females who often strike not in the forehead, but in the side. Males almost never: otherwise they would have killed each other in the first skirmishes. Ritual rules of fighting (of course, not consciously observed, but instinctive), developed over millions of years of evolution, are designed to protect fighters from severe injuries and death in skirmishes. This is amazing!
At first glance, duels between rams are quite dangerous: they run away and smash their heads together with a bang.
But they can afford this entertainment, because their horns, necks, and frontal bones are strong and can withstand such blows well. But the foreheads of goats are not suitable for ramming. They fight by striking their horns from above, and therefore stand on their hind legs before striking. You cannot keep a goat in the same enclosure with a ram. The goat is arrogant and does not calculate its strength well, while the ram has an armored skull. And if the ram, running away, hits the goat directly in the forehead, it can kill, break its neck or pierce its skull.
In addition to certain rules of fighting that limit injury, all animals and bovids also have special postures of submission and appeasement that allow the weak to avoid a fight. Thomson's gazelles are recumbent, with their necks extended along the ground. Some people fall to their knees. Therefore, the bull in the arena freezes and does not rush at the matador when he, kneeling right next to the bull’s muzzle, performs his tricks. The healthy instincts of an animal paralyze its aggressiveness, and a man with a sword, violating the morality of nature, acts in this case as a sadist: the continuation is well known to everyone.
That's all about the horns for now. Now about those who wear them on their heads.
This is an extensive family. All in it are ruminants, all artiodactyls: 128 species. They are divided in different ways and into different numbers of subfamilies. Let's take as an example a division that is perhaps the least complex:
1. Bovine: 13 wild and domesticated species of bovine (buffalo, zebu, gaur, guyal, cowrey, bison, bison, yak, etc.); 9 species of African antelopes (kudu, nyala, sitatunga, eland, bongo, etc.) and 2 species of Asian antelopes (nilgai and four-horned).
2. Duikers: the smallest of the antelopes, 17 species, all African.
3. Horse antelopes: waterbucks, ridbucks, oryxes, bases, saber-horned and horse antelopes, cow antelopes (topi, kongoni, wildebeest) - 24 species, all African, except for the Arabian oryx, which was almost exterminated.
4. Gazelles: impalas, dik-diks, oribi, beirs, gerenuk (giraffe gazelle), Thomson's gazelle, goitered gazelle, gazelle - 37 mainly African and partly Asian species.
5. Caprines: goats, rams, chamois, gorals, saigas, takins, musk oxen– 26 mainly Asian, European, partly North American and African species.
In South America there are no wild bovids, just like in Australia.
So, about the bulls. But before we begin, let’s digress a little for one necessary clarification. It concerns the word "antelope", which is more literary and everyday than strictly zoological scientific significance. In general, antelopes are usually called bovids that are not bulls, rams or goats. Medium-sized antelopes are also called gazelles, and the smallest ones are called duikers.
Greater kudu live in Africa - from Ethiopia to Angola and the Zambezi River in the south. The lesser kudu is found only in Somalia and eastern Africa.
Greater kudu live in Africa - from Ethiopia to Angola and the Zambezi River in the south.
The lesser kudu is found only in Somalia and eastern Africa.
“The beast is like a horse, terrible and invincible, with a large horn between its ears, its body is copper, and it has all its strength in the rose. Doesn't have any friends, lives 532 years. And when he throws his horn into the sea, and from it the worm grows; and from this there is a unicorn beast. But an old beast is not strong without a horn, becomes orphaned and dies.”
This is how the Russian alphabet books talked about the unicorn, they actually talked too “literarily”, because the prototype of the unicorn, as it turns out, was... a bull.
Archaeologists, excavating the sites of ancient cities of the Middle East, found Assyrian and Babylonian bas-reliefs and writings, from which it turned out that the Hebrew word “reem”, translated by the compilers of the Greek Bible as “unicorn”, actually meant a wild aurochs bull, completely two-horned.
The royal, or dwarf, antelope is the smallest of the antelopes: its height is only 25 - 30 centimeters. Her jumps are magnificent - almost three meters in length. Royal antelopes live in West Africa (Liberia, Nigeria). Second, somewhat more close-up view– in Nigeria and Cameroon.
So, the tour. He is up to two meters tall (at the withers) and weighs a ton! The color is black, the cows and calves are red. But you can argue about color... Remember the epics: “She wrapped Dobrynya in a bay aurochs”, “Where bays go nine aurochs”... Our ancestors were not color blind to confuse black with red! And yet, the tour is usually considered black, or rather, “he was black,” where the short “was” completely deprives us of the opportunity to know the true truth.
Because these bulls are no longer there. They were exterminated. And although this happened quite recently, the tour was thoroughly forgotten everywhere. It remains in epics, proverbs, some ancient rituals (for example, at Christmas time they dressed up as a tour) and in the names of places and surnames: Turovo, Tury, Turov log, Turova howl, Turzhets, Turov. The canton of Uri in Switzerland, of which Dostoevsky's Stavrogin was called a citizen, also owes its name to a wild bull: “Urus” in Latin, “ur” in German - the names of the tour.
But still, the assertion that the bull was black has serious grounds. Various images of the tour have reached us, and the best of them is the famous Augsburg painting. It was found in an antiquarian shop by the English zoologist Smith. It was drawn at the beginning of the 16th century by some Polish artist (and just about three hundred years ago the aurochs disappeared from the face of the Earth). This, it turns out, is a “posthumous” portrait (it disappeared, only a copy made by Smith survived) depicted the aurochs black - one must think, not for the sake of mourning.
But, of course, whatever it is, the image cannot serve as sufficiently serious evidence, because artists in all centuries were very prone to various liberties in their works (Assyrian and Babylonian bas-reliefs, for example, on which aurochs are one-horned, and horses are “double-legged” ": they only have two legs).
The proof is elsewhere. In 1921, German zoologists brothers Lutz and Heinz Heck, having traveled around Europe in search of “tur-like” bulls and cows (and finding suitable ones), began a remarkable experiment: using backcrossing methods, they decided to revive the aurochs.
The “restored” aurochs have everything like the extinct one: black color, large sharp horns. And the cows and calves are bay, which means that geneticists have achieved the most difficult thing: sexual and age dimorphism, that is, different colors and appearance of females, males and cubs. And finally: the “restored” tour is so similar to the one depicted in the Augsburg drawing that it seems as if it was drawn from it.
But back in the last century, even some serious naturalists did not believe that there was such a bull on Earth - an aurochs. Everything that the ancients told about him was attributed to the bison. Even V.I. Dal equates the words “tour” and “bison,” although he might not have done so, because by the time he was compiling his famous dictionary, the French anatomist and paleontologist Georges Cuvier had already proven that a long-horned bison once lived large bull - tour.
Duikers - there are probably seventeen species - are found throughout Africa south of Sudan. The height at the shoulders of different species ranges from 35 to 50 centimeters, and the weight ranges from 5 to 65 kilograms. All except the gray duiker, in which females are usually hornless, both sexes bear small horns.
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Sizes from small to large. Thus, Neotragus pygmaeus has a height at the withers of about 25 cm and a weight of 2-3 kg, and the bison has a height at the withers of up to 200 cm and a weight of up to 1000 kg. The general build ranges from light and slender to heavy and massive. The limbs are usually high. Males, and in many species also females, have a pair of unbranched horns (Tetracerus has two pairs). Horns are permanent, irreplaceable bony outgrowths of the frontal bones, covered on the outside with a horny sheath of epidermal origin. Antler growth, in contrast to deer, comes from its base. Thus, the top of the horn represents its oldest part. Characterized by periodic intensification and slowdown of horn growth, as a result of which peculiar rings are formed on its horny surface. The shape of the horns is extremely varied - from completely straight, long and thin to short, thick and strongly curved or spirally twisted. If the direction of bending or twisting of the horn occurs inward, towards the horn of the opposite side, then such horns are called homonymous, but if the right horn is folded or bent to the right, and the left one to the left - heteronymous. In cross-section, horns are round, oval or triangular. On their surface there are often protrusions, transverse folds and rings or longitudinal ribs.
The color is very diverse - from white to almost black, usually without sharp color patterns. Many species have a white field on the thighs - a “mirror”. The skin usually contains many specific glands: preorbital, interhorn, inguinal, interdigital, caudal and etc. 1-2 pairs of nipples.
There are 4 toes on the limbs (rarely 2), but the lateral toes (II and V) are greatly shortened and, although they have small hooves, they usually do not touch them when walking on hard ground. Only the proximal and distal parts of the metacarpal bones of the lateral fingers are preserved.
The frontal bones are strongly developed in the skull. The parietal bones are moved back. The lacrimal bone has a highly developed facial part with or without a fossa for the preorbital gland. Usually there is only one opening of the lacrimal canal. The ethmoidal foramina are absent or poorly developed. The bones of the skull are highly pneumatized. The premaxillary bone is usually relatively small, the maxillary bone is very large. Sometimes the second premolars in the lower, and occasionally in the upper jaws, do not develop or fall out early. The cheek teeth are hypselodont and tetraselenodont (four-lobed).
The stomach is complex, clearly divided into 4 sections: rumen, mesh, book and abomasum. Gallbladder usually available. The placenta is polycotton-iceous.
Distributed widely across the globe. The restored range covers Africa (without Madagascar), Europe (except for the British Isles), going north to the south of the Scandinavian Peninsula, the Gulf of Finland, the Upper Volga, Samara Luka and the south of the Urals. Beyond the Urals, the range includes the south of Western Siberia and most of Central and Eastern Siberia and the Far East. In the southeast and south of Asia, the range covers the entire southern part The Asian mainland with most of the adjacent islands. In the New World, the range occupies most of North America south to California, Florida and northern Mexico, the Arctic archipelago, northern and East Coast Greenland. As a result of excessive fishing or other reasons, the ranges of most species have been significantly reduced.
They live in a wide variety of places - from dense forests to steppes, semi-deserts and deserts on the plains, in the foothills and high mountains - higher than almost all other mammals (up to 5500 m above sea level). However, the largest number of species inhabit open spaces. They keep in herds, sometimes very large - up to several thousand heads. Much less common in small groups or alone. They feed on plants, mainly herbs.
Most species are polygamous, although some are monogamous. The males of some bovids have a harem of females during the breeding season. Inhabitants of the tropics, as a rule, have no seasonality in reproduction. The duration of pregnancy is 4-11 months. There are from one to 4-5 cubs in a litter.
Many species of bovids are of significant importance as game animals, from which they obtain meat and leather. A number of species served as the ancestors of the most important domestic animals.
The bovid family contains 140 species, ranging from the 5-kg dik-dik to the 1,000-kg bison. An important difference are horns: there is almost always one pair of them (the exception is the genus of four-horned antelopes), and the length can range from 2 cm to 1.5 meters. Some species have horns only on males, but most have horns on both sexes. These are bony structures firmly connected to the skull. Unlike deer and pronghorn, bovids never have branched antlers.
The largest representative of the family is the gaur (up to 2.2 m tall at the withers and weighing more than a ton), and the smallest is the dwarf antelope (weighs no more than 3 kg and is as tall as a large domestic cat).
The majority of bovids live in open areas. African savannas provide an ideal living space for many species. There are also species that live in mountainous areas or forests.
Most members of the family are herbivores, although some antelopes can also eat animal food. Like other ruminants, bovids have a four-chambered stomach, which allows them to digest plant foods, such as grasses, that cannot be used as food by many other animals. Such food contains a lot of cellulose, and not all animals are able to digest it. However digestive system ruminants, which are all bovids, are able to digest such food.
The horns are attached to the protruding frontal bone. The length and width are different (the girth of argali horns, for example, is 50 cm). The horns of bovids grow throughout their lives, but never branch. They consist of a substance of epidermal origin. Mainly, horns are used by males in skirmishes with relatives.
Historically, bovids are a relatively young group of animals. The most ancient fossils that can be confidently classified as bovids are the genus Eotragus (English) Russian from the Miocene. These animals resembled modern crested duikers, were no larger than roe deer and had very small horns. Even during the Miocene, this genus was divided, and in the Pleistocene all important lines modern bovids. In the Pleistocene, bovids migrated along the then existing natural bridge from
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